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I think the difference depends mainly on your aero and rolling resistance. The less resistance you have one the car, the more you can take advantage of hills with less inclination. I've noticed this especially with my recent wheel swap.
Part of the issue I think with slight downgrades that you can't EOC on is that even if it would only take 1hp to maintain speed the car needs to put out like 4 or 5 hp just to overcome all the losses in the engine and transmission, which lowers potential mileage around those sorts of hills.
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