My vote is to haul the bike in the bed. It would seem to me that's the choice with the best aero.
Up front seems like it would be pretty aero dirty, but I really like that setup when you trailer. By the way thats a very cool little 5th wheel kamper!
I think a decent amount of the side profile of the bike would still be in the wind above the tailgate if it's on the rear hitch, maybe I'm wrong here, not sure of the carrier/bike height in relation to the tailgate. But if a decent portion of the side of the bike is in play it would seem to me to be kinda dragy.
I think with it in the bed and strapped down real tight (To pull it as low as possible as well as keep it secure.) very little of it will be left in the direct wind, maybe even swing the mirrors down or spin them off. I'm not a big fan of loading/unloading bikes into beds solo so thats something to consider also. With a wide enough ramp it is better but still not fun.
If you have the time before your trip maybe you could do some coast down testing to see which location gives the best results.
What kind of MPG do you see while towing with the pictured setup? I will be seeing my first real highway results with my 6.8 liter V-10 Excursion towing a 28' 9k# Toyhauler in April, 1200+ miles. I will be keeping the speed down, DWL'ing and watching my Ultra-Gauge to make the most of a thirsty situation.
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