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Welcome to the site.
Windows up/down really depends on speed. If you're going 25 mph windows down is probably better. However, at higher speeds windows down and A/C on is probably going to be better. Obviously windows up and fans on full speed is the best, but if heat doesn't allow for it I usually just turn the A/C on, temperature all the way to cold, and keep the fan speeds as low as is comfortable.
Highway rpm really isn't adjustable, so the general rule is slower is better.
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