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I think it's because the contact patch of the tire between the road deforms the roundness of the tire. That deformation becomes rolling resistance and has been described like your driving up a very shallow grade. The larger diameter tire 'should' rotate into the contact patch at a shallower angle and thus deflects less as it deforms into the contact patch.
I would love to have a tire with 0 aspect ratio and no deflection. Almost like a solid metal train wheel which has a rolling resistance equal to or less than the best racing bicycle tires.
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I try to be helpful. I'm not an expert.
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