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Wheel/Tire weight
I know for my 1986 Jeep, the studded winter tires weigh in at 37lbs each and I get 20-22 miles to the gallon (in the winter). My summer tires weigh in at 28lbs each and I average 26+ miles to the gallon with them. The warmer temps and type of rubber/tread design also play a role but I do not know the math behind it all. I bought my summer tires and rims from tire rack because I could look up the weight of the rim and tire before I bought them.
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