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Each wheel-tire assembly is in essence a flywheel. It takes energy to spin up all 4 wheels including all rotating mass (brake rotors-drums). It will improve you mileage but with colder temps, as you posted, the benefit will be offset by loses due to temperature.
Greater rotating mass requires more power to get to the same speed but coasting will suffer slightly. Overall you should see an increase in mileage. I would keep the lighter wheels due to the better road feel, steering response and ride quality. Ride quality especially with higher inflation pressures.
regards
mech
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