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IMHO most of the action of combustion takes place early on; perhaps whether an engine is square or not is of little consequence, or less consequence than simply what the stroke is because the stroke defines what the piston speed will be and from what I can tell optimal piston speed is 1000-1200 ft/mn. Of course with a longer stroke you get your 1000-1200 at a lower rpm with all the good things that brings.
Perhaps the high rotating mass is helping with lug-a-bility- short shifts, tall gears, and whatnot. Spool it up and it can get moving with those long legs without stalling
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