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That section on wheel flow in Hucho also has a very helpful graphic visualizing airflow around a spinning wheel. Three pairs of vortices are shed from the wheel--two from the top, two from the bottom, two from the center--all spinning inward because of the rounded shoulders, and all raising the pressure behind the wheel due to entrainment.
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Can you provide a quote? In the picture at Permalink #15, you can see the front wheel's bottom vortexes passing the rear tire. The dirt carried over-top in the tread is well contained. I'd wondered about those streaks coming off from mid-wheel.
Another possibility on the spat is that it is to cover the tire when it is turned. Best practice in the rear is a fence right up against the inside of the tire.
