To answer your earlier question about how much air flow is needed for a 125,000btu furnace... 1600cfm. And the airflow required for a 5 ton a/c is 2000cfm.... (and..17x20" duct is about 1600cfm).
So yes, your biggest gain with your current equipment would be to fill that 17x20" with as much air as possible. I think it would be very easy to open up the return grille to be large enough...(even going a bit bigger, say 2 20x30" return grilles.)
It also looks like you could tap the right side of the return just behind the grille to penatrate the bedroom wall, adding a return to each bedroom is recommended, It looks like you have high ceilings (with fresh paint ).ducting some return air from up high would be good if you didn't mind tearing out drywall....a standard 2x4 wall cavity lined with a 3.5x14" duct is only good for 100cfm.
Is it also possible to run a other 12" round return into the other floor somewhere?
Now I'm curious about the number of and sizes of your supply vent registers....
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