Frictional losses are most definitely the biggest culprit for engine braking... Imagine it this way -
You've got a 300 lb person sitting on your chest, and you're flat on your back on a hardwood floor, with your mouth and nose covered by a slightly porous wet rag. You're laboring to breathe at this point...
Remove that rag, but leave the fat guy. Now you can breathe freely, but your chest has trouble rising to get an "easy" breath...
When you open the throttle, you're just taking the rag off your engine's face - get it?
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