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negative boost is negative boost, regardless of the engine type. The only difference is the carbed cars don't dynamically adjust fueling, that's all.
All this study is saying in the end is that even if your filter is clogged, the negative boost is never enough to have a significant impact on fuel economy when driven in normal conditions. A free flowing filter is always better, but so marginally better it's insignificant. Ram air reduces, and can even negate, negative boost, but again it's not much significant under normal driving conditions. WAI has ramifications in the combustion process itself so it's not just about negative boost in that case.
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