How are you measuring your fuel consumption? (Instrumentation?)
It's possible you're so far out of the ideal BSFC zone for that engine at low RPM in the highest gear that fuel consumption takes a hit (vs. same gear, higher RPM).
I've yet to see that in any instrumented car I've driven though, where the "lowest RPM in the highest gear" rule of thumb always seems to return the best fuel economy.
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