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Drivetrain-wise, the low hanging fruit for highway driving is in the gearing. Having a cam that doesn't have a ton of overlap (e.g. a cam not for top end power) is generally going to deliver better economy too.
But as brucepick says, you'll likely get more mileage out of even subtle body changes. Smooth underbody paneling, wind deflectors in front of the wheels, maybe an air dam, fenders that close off the sides of the wheel well gap, maybe even some kind of diffuser for the rear.
Nice looking Jeep.
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