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The MPGuino will indeed track mileage weather the car is on or off. When it is wired, it is wired into a constant 12V source, so it never turns off (only the backlight turns on and off).
Edit: I should clarify. The MPGuino will track mileage weather the car is on or off. However, if the car is off (key in the OFF position), there is no voltage going to the speed sensor, so the guino gets no signal and thus can not track mileage. As soon as you turn the key back to on, the sensor has voltage and the guino continues to log. This can all be avoided with a kill switch though.
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